Charis

Charis is a genus of the Riodinini tribe of metalmark butterflies (family Riodinidae). Nineteen species have been identified within the Charis cleonus complex (however, further analysis has revised this clade to contain 22 species) and eight species within the Charis gynaea group (clade). Charis butterflies are common in the Neotropics and often live in primary and secondary growth. The Charis cleonus group exhibits contemporary parapatric distributions throughout Amazonia and are thought to have speciated allopatrically; residing in "areas of endemism". Some evidence suggests that Charis (among other Riodininae groups) are reproductively isolated by mating preferences for different topographic areas and different times—of which, may have promoted speciation between the various groups. Species Charis cleonus complex Charis major subgroup Charis ma Harvey & Hall, 2002 Charis major (Lathy, 1932) Charis matic Harvey & Hall, 2002 Charis cleonus subgroup Charis ariquemes Harvey & Hall, 2002 Charis brasilia Harvey & Hall, 2002 Charis breves Harvey & Hall, 2002 Charis cacaulandia Harvey & Hall, 2002 Charis caryatis Hewitson, 1866 Charis cleonus (Stoll, 1781) Charis cuiaba Harvey & Hall, 2002 Charis humaita Harvey & Hall, 2002 Charis ipiranga Harvey & Hall, 2002 Charis iquitos Harvey & Hall, 2002 Charis manicore Harvey & Hall, 2002 Charis manu Harvey & Hall, 2002 Charis maues Harvey & Hall, 2002 Charis negro Harvey & Hall, 2002 Charis palcazu Harvey & Hall, 2002 Charis rocana Harvey & Hall, 2002 Charis santarem Harvey & Hall, 2002 Charis tapajos Harvey & Hall, 2002 Charis tefe Harvey & Hall, 2002 Charis gynaea group Charis barnesi Hall & Harvey, 2001 Charis callaghani Hall & Harvey, 2001 Charis gallardi Hall & Harvey, 2001 Charis gynaea (Godart, 1824) Charis nicolayi Hall & Harvey, 2001 Charis smalli Hall & Harvey, 2001 Charis zama Bates, 1868 References Riodininae